Brendan Fletcher

2003. As a parasitic fungal outbreak begins to ravage the country and the world, Joel Miller attempts to escape the escalating chaos with his daughter and brother. Twenty years later, a now hardened Joel and his partner Tess fight to survive under a totalitarian regime, while the insurgent Fireflies harbor a teenage girl with a unique gift.

While the Governor shuts down hunting season, Joe searches for truth in the aftermath of the new murders at the crime scene. Meanwhile, Marybeth digs deep into Stop the Slaughter and Klamath Moore and discovers a disturbing pattern of events. Nate and Cricket search for answers -- and safety while on the run. Missy re-examines her relationship with McLanahan.

Basilone's celebrity grows as he travels across the country on the war bonds tour. On Pavuvu, Sledge, assigned to the 5th Marines, 1st Marine Division, is briefly reunited with Phillips and Leckie rejoins his company. Sledge then gets his first taste of combat as he, Leckie, and the rest of the 1st Marine Division meet fierce Japanese resistance while landing on the intricately and heavily defended coral island of Peleliu.

Despite the suffocating 115-degree heat and a lack of clean drinking water, Sledge, Leckie and the other Marines confront the highly-fortified enemy as they attempt to capture the Peleliu airfield. After seeing his comrades badly injured, Leckie himself is wounded and evacuated from the island. Sledge witnesses the shocking truth about what is sometimes required to survive and fight another day.

The Marines, including the newly christened Sledgehammer, continue the battle of Peleliu against an enemy determined to fight to the last man. Devastated by the loss of a revered leader, and witnessing unimaginable barbarity on both sides, Sledge veers to the very edge of moral collapse. Their objective finally secured, the Marines return to Pavuvu fundamentally changed by their experience on Peleliu.